Where does “extranucleolar” come from?

extranucleolar (English) comes from English extra-, from Latin extrā, from Latin exter, from Proto-Italic *eksteros.

extranucleolar (English): Outside of a nucleolus

Definitions

  1. Outside of a nucleolus

Ancestry of “extranucleolar”, step by step

StepLanguageWordMeaning
1Englishextra-Outside of, beyond
2Latinextrāon the outside
3LatinexterOn the outside, outward, external, outer; Of...
4Proto-Italic*eksteros
Every word from Proto-Italic *eksterosEvery word from Latin exterEvery word from Latin extrā