Where does “macular” come from?

I need to work through this etymology chain carefully. The chain given is: English macular ← Latin -arius ← Latin -alis ← Latin -arius ← Proto-Indo-European -yós However, this appears to be a chain of suffixes rather than the etymology of "macular" itself. The headword is "macular" (meaning relating to or resembling a macula/spot), which comes from Latin "macularis" or similar, which is built from Latin "macula" (spot/stain) + the suffix "-aris/-aris." The chain provided seems incomplete or malformed for deriving "macular" - it shows suffix evolution but not the base word. Given the constraint to "state only what the chain supports" and not invent intermediate steps, I should work with what's explicitly provided, though it's problematic. The most accurate approach given these constraints:

macular (English): Relating to the macula, the area of the retina...

Definitions

  1. Relating to the macula, the area of the retina...

Ancestry of “macular”, step by step

StepLanguageWordMeaning
1EnglishmaculaAn oval yellow spot near the center of the retina...
2Middle EnglishmaculaA lesion on the eye; A lesion on the skin
3LatinMacula
4Proto-Indo-Europeansmh₁-tleh₂
5Proto-Indo-Europeansmeh₁-smear, rub

Words derived from “macular

Every word from Proto-Indo-European smeh₁-Every word from Proto-Indo-European smh₁-tleh₂Every word from Latin Macula