Where does “quadrinucleated” come from?

quadrinucleated (English) comes from English quadri-, from Latin quattŭor, from Portuguese quarto, from Old Portuguese quarto, from Latin quārtus, from Proto-Indo-European kʷeturtos, from Proto-Indo-European kʷetwóres, from Proto-Indo-European kʷe- — and.

Ancestry of “quadrinucleated”, step by step

StepLanguageWordMeaning
1Englishquadri-API; Four
2Latinquattŭorfour; 4
3Portuguesequartoordinal form of quatro; fourth; quarter; fourth;...
4Old Portuguesequartoquarter
5Latinquārtusfourth
6Proto-Indo-Europeankʷeturtos—
7Proto-Indo-Europeankʷetwóresfour
8Proto-Indo-Europeankʷe-and
Every word from Proto-Indo-European kʷe- →Every word from Proto-Indo-European kʷetwóres →Every word from Proto-Indo-European kʷeturtos →