Where does “unguligradus” come from?

unguligradus (Late Latin) comes from Latin gradus, from Proto-Italic graðus, from Proto-Indo-European gʰradʰ-.

Ancestry of “unguligradus”, step by step

unguligradus traces back through two separate lines of ancestry.

via Latin gradus

StepLanguageWordMeaning
1Latingradusa step, pace; a stage, degree; a rank
2Proto-Italicgraðus
3Proto-Indo-Europeangʰradʰ-

via Latin ungula

StepLanguageWordMeaning
1Latinungulahoof, claw; a horse; an aromatic spice
2Latinunguisfingernail, toenail; claw; hoof
3Proto-Indo-Europeanh₃negʰ-nail
Every word from Proto-Indo-European gʰradʰ-Every word from Proto-Italic graðusEvery word from Latin gradus