Where does “maculandus” come from?

maculandus (Latin) comes from Latin maculō, from Latin Macula, from Proto-Indo-European smh₁-tleh₂, from Proto-Indo-European smeh₁- — smear, rub.

Definitions

Ancestry of “maculandus”, step by step

StepLanguageWordMeaning
1Latinmaculōto stain, make spotted
2LatinMacula—
3Proto-Indo-Europeansmh₁-tleh₂—
4Proto-Indo-Europeansmeh₁-smear, rub
Every word from Proto-Indo-European smeh₁- →Every word from Proto-Indo-European smh₁-tleh₂ →Every word from Latin Macula →