Where does “maculans” come from?

maculans (Latin) comes from Latin maculō, from Latin Macula, from Proto-Indo-European smh₁-tleh₂, from Proto-Indo-European smeh₁- — smear, rub.

maculans (Latin): staining, spotting; defiling, polluting;...

Definitions

Step	Language	Word	Meaning
1	Latin	maculō	—
2	Latin	Macula	—
3	Proto-Indo-European	smh₁-tleh₂	—
4	Proto-Indo-European	smeh₁-	smear, rub

maculans — Etymology, Origin & Word History | Etymology Explorer