Where does “*buto” come from?

*buto (Proto-South Sulawesi) comes from Proto-Malayo-Polynesian butuq, from Proto-Austronesian buCuq — testicles.

Ancestry of “*buto”, step by step

StepLanguageWordMeaning
1Proto-Malayo-Polynesianbutuqtesticles
2Proto-AustronesianbuCuqtesticles

Words derived from “*buto”

Every word from Proto-Austronesian buCuq →Every word from Proto-Malayo-Polynesian butuq →